This recent study (see link below) has a control group which the other study concluding uphill interval improved 5km times did not. Because the study linked below has a control group, their conclusions is supported that,"both uphill and level-grade interval training can induce significant
improvements in a run-to-exhaustion test in well-trained runners at the
speed associated with VO2max, but that traditional level-grade training
produces greater gains." I believe it would be safe to say that these types of training would lead to an improved 5km time.
http://www.ncbi.nlm.nih.gov/pubmed/22996027
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